What it means in practice
In a rights of nature framework, the rights-holder is the “entity” whose interests are represented. The law usually names (or describes) the rights-holder and clarifies its scope (for example, whether it includes tributaries, wetlands, and connected habitat).
Why it matters for rights of nature
Identifying a rights-holder shifts the focus from nature as property or a resource to nature as a legal interest the system must hear and protect. It also makes enforcement workable: if there is a rights-holder, then there can be representation, violations, and remedies.
See also
Natural community / ecosystem; Legally recognized rights; Guardian; Guardianship; Rights of Nature